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Innate human propensities for cooperation with strangers, shaped during the Pleistocene in response to rapidly changing environments, could have provided highly adaptive social instincts that more recently coevolved with cultural institutions; although the biological capacity for primate sociality evolved genetically, the authors propose that channeling of tribal instincts via symbol systems has involved a cultural transmission and selection that continues the evolution of cooperative human capacities at a cultural rather than genetic level - and pace.