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One of these groups can be traced as far back as 35,000 years, thanks to a skeleton discovered at a site in Belgium called Goyet. The Goyet-related people spread across Europe, only to be replaced on much of the continent near the end of the Ice Age by a genetically distinct population. The earliest sign of the second group appears 14,000 years ago, known to researchers by DNA in a skeleton at an Italian site called Villabruna.

the Goyet and Villabruna people coexisted. Hunter-gatherers across the peninsula had a mixture of ancestry from the two peoples.

But whatever solitude Iberia might have offered came to an end about 7,500 years ago, when new people arrived with crops and livestock. These first farmers, originally from Anatolia, brought with them a distinctive genetic signature.

Ninety percent of the DNA from the later skeletons derives from the Anatolian farmers; 10 percent comes from the hunter-gatherers.

Starting about 6,000 years ago, Dr. Olalde and his colleagues found, hunter-gatherer ancestry in Iberian farmers actually increased to 20 percent.

A skeleton from an elaborate grave in central Spain about 4,400 years old belonged to a man whose ancestry was 100 percent North African

findings suggest that people were moving into Iberia from Africa more than 3,000 years before the rise of the Roman Empire. “These are cosmopolitan places,”

About 4,500 years ago, still another wave of people arrived, profoundly altering the makeup of Iberia.A few centuries earlier, nomads from the steppes of what is now Russia turned up in Eastern Europe with horses and wagons. They spread across the continent, giving up nomadic life and intermarrying with European farmers.

DNA from the men, however, all traced back to the steppes. The Y chromosomes from the male farmers disappeared from the gene pool.

When they finally reached Iberia, these people spread out far and wide. “They really have an impact on the whole peninsula,”

To archaeologists, the shift is a puzzle.“I cannot say what it is,

he ruled out wars or massacres as the cause. “It’s not a particularly violent time,”

Iberian farmers originally lived in egalitarian societies, storing their wealth together and burying their dead in group graves.

over several centuries, palaces and fortresses began to rise, and power became concentrated in the hands of a few. Dr. Risch speculated that the cultural shift had something to do with the genetic shift

The Bronze Age in Iberia was followed by the Iron Age about 2,800

Iron Age Iberians could trace some of their ancestry to new waves of people arriving from northern and Central Europe, possibly marking the rise of so-called Celtiberian culture on the peninsula.

scientists found a growing amount of North African ancestry in skeletons from the Iron Age. That may reflect trade around the Mediterranean, which brought North Africans to Iberian towns, where they settled down.

North African ancestry increased in Iberia even more after Romans took control. Now the peninsula was part of an empire that thrived on widespread trade. At the same time, people from southern Europe and the Near East also began leaving an imprint

The Basque speak a language that is unrelated to other European tongues. Some researchers have speculated that they descended from a population that had been distinct since the Bronze Age or earlier.Genetically, at least, that doesn’t seem to be the case. Before the Roman era, the Basque had DNA that was indistinguishable from that of other Iron Age Iberians

After the fall of Rome, ancient DNA in Iberia reflects its medieval history. Skeletons from the Muslim era show growing ancestry from both North Africa and sub-Saharan Africa.

The researchers were also able to group Spaniards into five genetic clusters. On a map, these groups form five strips running north to south. Those strips line up neatly with history.

The victory of John I may be regarded as a triumph of the national spirit over the feudal attachment to established order. Because much of the older nobility sided with Castile, John rewarded his followers at their expense and the crown’s. Meanwhile, commerce prospered, and the marriage of John’s daughter Isabella to Philip III (the Good) of Burgundy was to be followed by the growth of close trading relations between Portugal and Philip’s county of Flanders.

In 1437, during the short reign of John’s eldest son, Edward (Duarte; 1433–38), an unsuccessful attempt to conquer Tangier was made by John’s third son, Prince Henry the Navigator, and his younger brother Ferdinand (who was captured by the Moors and died, still unransomed, in 1443). Edward’s son Afonso V (1438–81) was still a child when Edward died, and Edward’s brother Pedro, duke of Coimbra (Dom Pedro), had himself made regent (1440) instead of the widow, Leonor of Aragon.

Having married Joan, daughter of Henry IV of Castile, Afonso laid claim to the Castilian throne and became involved in a lengthy struggle with Ferdinand and Isabella in the region of Zamora and Toro, where he was defeated in 1476. He then sailed to France in a failed attempt to enlist the support of Louis XI, and on his return he concluded with Castile the Treaty of Alcáçovas (1479), abandoning the claims of his wife. Afonso never recovered from his reverse, and during his last years his son John administered the kingdom.

John II (1481–95) was as cautious, firm, and jealous of royal power as his father had been openhanded and negligent.

his forces departed in June 1578 and on August 4 were utterly destroyed by the Moors in the Battle of the Three Kings near Alcazarquivir (Ksar el-Kebir). Sebastian and some 8,000 of his forces were killed, some 15,000 were captured, and only a handful escaped.

As a condition of his marriage to Isabella, Manuel was required to “purify” Portugal of Jews. After Jews were expelled from Spain in 1492, John II had admitted many Jewish refugees; he had taxed the Jews heavily but was also to supply ships for them to leave Portugal.

If Manuel failed to realize his dream of ruling Spain, his son John III (1521–57) lacked the power to resist Castilian influence.

In 1529 the settlement by the Treaty of Zaragoza (Saragossa) of a dispute over the possession of the Moluccas (an island group part of present-day Indonesia) removed an obstacle to Portuguese-Spanish understanding, and the line dividing Portuguese and Spanish interests in the New World (established by the Treaty of Tordesillas) was matched by a similar line in the Pacific.

John III was succeeded by his grandson Sebastian (1557–78), then only three years old. As a child Sebastian became obsessed with the idea of a Crusade against Morocco.

Predeceased by his legitimate son, John II was succeeded by his cousin the duke of Beja, as Manuel I (1495–1521), known as “the Fortunate.” Manuel, who assumed the title of “Lord of the Conquest, Navigation, and Commerce of India, Ethiopia, Arabia, and Persia,” inherited, because of the work of John II, a firmly established autocratic monarchy and a rapidly expanding overseas empire. Drawn toward Spain by the common need to defend their overseas interests as defined by the Treaty of Tordesillas (1494), Manuel nourished the hope that the whole peninsula could be united under the house of Aviz; to that end he married Isabella, eldest daughter of Ferdinand and Isabella. However, she died in 1498 while giving birth to a son, Miguel da Paz. This child, recognized as heir to Portugal, Castile, and Aragon, died in infancy. Manuel then married Isabella’s sister Maria (died 1517) and eventually Eleanor, sister of the emperor Charles V.

Sebastian was succeeded by his great-uncle, Cardinal Henry (1578–80), a brother of John III. His age and celibacy made it certain that the Portuguese throne would soon pass from the direct line of Aviz. Philip II of Spain, nephew of John III and husband (by his first marriage) of John’s daughter Maria

They coexisted in Europe for thousands of years, apparently trading tools and sharing cultures. Some groups survived the Ice Age, while others vanished,

when farmers arrived in Europe about 8,000 years ago, they encountered the descendants of this long history, with light-skinned, dark-eyed people to the east, and possibly dark-skinned and blue-eyed people to the west.

“We lack still an understanding of why these movements were triggered. What happened here, why it happened — it’s strange.”

Modern humans arose in Africa and expanded to other continents about 60,000 years ago

These early Europeans have almost no genetic link to younger remains of hunter-gatherers. It appears that the first modern humans in Europe may have disappeared along with the Neanderthals

he oldest DNA of modern humans in Europe, dating back 45,000 years, undermines such a simple story. It comes from people who belonged to a lost branch of the human family tree. Their ancestors were part of the expansion out of Africa, but they split off on their own before the ancestors of living Europeans and Asians split apart.

About 33,000 years ago, as the climate turned cold, a new culture called the Gravettian arose across Europe. Gravettian hunters made spears to kill woolly mammoths and other big game. They also made so-called Venus figurines that might have represented fertility.

When the glaciers retreated, some descendants of the Fournol continued living in Iberia. But others expanded north as a new population, which Dr. Posth and his colleagues called GoyetQ2. “It really seems like a peopling of Europe after the last glacial maximum,

Dr. Posth and his colleagues found DNA in Gravettian remains scattered across Europe. The scientists had expected all of the individuals to have come from the same genetic population, but instead found two distinct groups: one in France and Spain, and another in Italy, the Czech Republic and Germany.

“They were very distinct, and this was a very big surprise to us because they practiced the same archaeological culture,”

Dr. Posth and his colleagues named the western population the Fournol people, and found a genetic link between this group and 35,000-year-old Aurignacian remains in Belgium.

They called the eastern group Vestonice, and discovered that they share an ancestry with 34,000-year-old hunter-gatherers who lived in Russia.

That genetic gulf led Dr. Posth and his colleagues to argue that the Fournol and Vestonice belonged to two waves that migrated into Europe separately. After they arrived, they lived for several thousand years sharing the Gravettian culture but remaining genetically distinct.

It’s clear from the new study that they were not isolated entirely from each other. In Belgium, the scientists found 30,000-year-old remains with a mix of Fournol and Vestonice ancestry.

About 26,000 years ago, the two groups faced a new threat to their survival: an advancing wall of glaciers. During the Ice Age, from 26,000 to 19,000 years ago, European hunter-gatherers were shut out of much of the continent, surviving only in southern refuges.

the refuge of the Iberian Peninsula, the region now occupied by Spain and Portugal, by studying DNA in the teeth of a 23,000-year-old man found in a cave in southern Spain. His DNA revealed that he belonged to the Fournol people who lived in Iberia before the Ice Age. The researchers also found genetic markers linking him to a 45,000-year-old skeleton discovered in Bulgaria.

When these groups arrived in Europe, Neanderthals had already been living across the continent for more than 100,000 years. The Neanderthals disappeared about 40,000 years ago, perhaps because modern humans outcompeted them with superior tools.

The Vestonice, by contrast, did not survive the Ice Age. When the glaciers were at their most expansive, the Vestonice may have endured for a time in Italy. But Dr. Posth and his colleagues found no Vestonice ancestry in Europeans after the Ice Age. Instead, they discovered a population of hunter-gatherers that appeared to have expanded from the Balkans, known as the Villabruna. They moved into Italy and replaced the Vestonice.

For several thousand years, the Villabruna were limited to southern Europe. Then, 14,000 years ago, they crossed the Alps and encountered the GoyetQ2 people to the north. A new population emerged, its ancestry three parts Villabruna to one part GoyetQ2.

This new people, which Dr. Posth and his colleagues called Oberkassel, expanded across much of Europe, replacing the old GoyetQ2 population.

another climate shift could explain this new wave. About 14,000 years ago, a pulse of strong warming produced forests across much of Europe. The Oberkassel people may have been better at hunting in forests, whereas the GoyetQ2 retreated with the shrinking steppes.

To the east, the Oberkassel ran into a new group of hunter-gatherers, who probably arrived from Russia. The scientists named this group’s descendants, who lived in Ukraine and surrounding regions, the Sidelkino.

in Iberia, there were no great sweeps of newcomers replacing older peoples. The Iberians after the Ice Age still carried a great deal of ancestry from the Fournol people who had arrived there thousands of years before the glaciers advanced. The Villabruna people moved into northern Spain, but added their DNA to the mix rather than replacing those who were there before.

When the first farmers arrived in Europe from Turkey about 8,000 years ago, three large groups of hunter-gatherers thrived across Europe: the Iberians, the Oberkassel and the Sidelkino. Living Europeans carry some of their gene

The Sidelkino people in the east had genes associated with dark eyes and light skin. The Oberkassel in the west, in contrast, probably had blue eyes and may have had dark skin

These three groups of hunter-gatherers remained isolated from each other for about 6,000 years, until the farmers from Turkey arrived. After this advent of agriculture, the three groups began mixing, the scientists found. It’s possible that the spread of farmland forced them to move to the margins of Europe to survive. But over time, they were absorbed into the agricultural communities that surrounded them.

every continent will likely have its own history of hunter-gatherer migrations.

it is now possible to extract human DNA from cave sediments rather than searching for bones and teeth.

In Castile an influential party of magnates, led by Alfonso Carillo, archbishop of Toledo (who later reversed himself), and opposed to King Henry IV, supported the succession claims of the princess Isabella, the king’s half sister, against those of his daughter, Joan.

It needed a forged papal dispensation for the marriage, the blackmailing of Henry IV into (wrongly) denying the paternity of his daughter (Joan), and, finally, several years of bitter civil war before Ferdinand and Isabella defeated Joan’s Castilian supporters and her husband, Afonso V of Portugal.

Ferdinand and Isabella ruled jointly in both kingdoms and were known as the Catholic Monarchs

In Spain the Catholic Monarchs had no formal right of exequatur, but they and their Habsburg successors behaved very much as if they did.

Some researchers believe that the introduction of seafood into the diet of early modern humans helped their cognitive development because of Omega-3 fatty acids and other brain-boosting nutrients. That contributed to cultural and technological developments that led them to migrate out of Africa and spread across the globe.

Zilhão published a study two years ago about 65,000-year-old cave paintings found in three caves on the Iberian Peninsula that are credited to Neanderthals. This aligns with another discovery of pendants and shells colored with pigments, also thought to be the work of Neanderthals.

Earlier this year, a separate analysis of clam shells and volcanic rocks from an Italian cave shows that Neanderthals collected shells and pumice from beaches. And due to specific indicators on some of the shells, the researchers also believe Neanderthals waded and dove into the ocean to retrieve shells, meaning they may have been able to swim.

One of the oldest genomes studied came from a thigh bone discovered in Goyet Cave in Belgium and given the unwieldy name GoyetQ116-1. Radiocarbon dating pegs the Goyet individual at some 35,000 years old,

Around 1,000 years after the Goyet individual was found, a new culture swept through Europe: the Gravettians. Analysis of genetic material from the time shows that art and artifacts weren't the only things changing. The Gravettians' DNA was significantly different from their Aurignacian predecessors, suggesting that they were a completely separate lineage.

Goyet guy's descendants retreated to the Iberian Peninsula (modern day Spain and Portugal) and waited for their time to come again.

It did, some 15,000 years later. Probably spurred by climate changes as glaciers began to recede, this dormant lineage expanded back into the rest of Europe, bearing a new culture known as Magdalenian.

Not long after that, their genomes started to look like those of people from the Middle East and the Caucasus, suggesting that new arrivals from the southeast were mingling with — and in some cases supplanting — the existing population.

This was a surprise, because researchers used to think that transition happened much later, when Turkish farmers introduced agriculture to Europe some 8,500 years ago.

The genetic analysis allowed researchers to trace the inexorable decline of Neanderthal DNA, which was two to three times more prominent in early human genomes than it is in modern-day ones. This supports theories that early humans interbred with Neanderthals, but that their DNA was toxic to us and gradually weeded out by natural selection over the course of millennia.

Numerous digs, most notably in Austria and Switzerland, have traced the outlines of the Celts. The artifacts offer evidence going back as far as about 800 B.C. The ancient Greeks and Romans also left written accounts of the Celts, and probably knew them well — the Celts sacked Rome around 390 B.C. and attacked Delphi in Greece in 279 B.C.

It seemed plausible that this group that had invaded Rome had invaded Ireland as well, and in the standard view, it was this people that eventually made it to Ireland.

For decades, however, archaeologists and other scholars have noted just how flimsy the evidence is for that standard account and how broad, nonetheless, is the application of the word.

n recent years, some archaeologists have proposed that the traditional story of the Celts' invasion was, in a sense, exactly wrong — the culture was not imported but exported — originating on the western edge of Europe much earlier than previously thought and spreading into the continent.

it flowed from the western edge of Europe, what he calls “the Atlantic zone” — into the rest of the continent. In many places of the Atlantic zone, he notes, people were buried in passages aligned with the solstices, a sign that they shared a unified belief system.

“From about 5,000 B.C. onwards, complicated ideas of status, art, cosmology were being disseminated along the Atlantic seaways,” Cunliffe said, and that culture then spread eastward.

“If we’re right, the roots of what is known as ‘Celtic’ culture go way way back in time,” Cunliffe said. “And the genetic evidence is going to be an absolute game-changer.”

If the new scholarship proves correct, exactly what to do with the word Celtic will probably be a matter of some dispute: Should it be applied to languages or cultures that, no matter how clearly defined, were largely uninfluenced by the historical Celts of continental Europe?

Complicating any answer are old ethnic antagonisms: The old notions of a distinct “Celtic race” or “Irish race” have been used not just for poetic tributes, but for scorn.

According to the genetic research, the Irish are at the extreme end of a genetic wave that washed across Europe, a wave of migrants that swept westward from above the Black Sea across Europe about 2,500 B.C.

The way to think about genetic variation in Europe is that it is more of a gradient than it is of sharp boundaries," said Bradley, the DNA researcher. "Sometimes, cultural features like language and natural borders can coincide with genetics, but most times not. Genetics is fuzzy, and it doesn't follow political and cultural borders."

"The public will always want a place on the map and for someone to point and say, 'This where the Irish are from,'

"But there’s going to be no way to do that. These groups were frequently traveling east-west across Europe, from one place to another. Everyone is a mix.”

The castle presented a formidable problem; the ships’ guns could not be elevated sufficiently to sweep the summit of the rock, and it was not until a number of bomb vessels had been summoned that the fortress could be brought under effective fire

resolutely determined to retain their Bourbon monarch, Philip V, grandson of Louis XIV, on the throne willed to him in 1700 by the last of the Spanish Habsburgs

The Earl of Galway’s army received a sharp defeat at Almanza some sixty miles inland from Alicante on April 25 th, 1707, and within the next twenty months the Bourbonists had recaptured the important cities of Lerida and Tortosa besides many more minor places

On the Spanish mainland, however, the forces of Philip V undoubtedly enjoyed the main initiative from Almanza onwards, while the Allied generals spent much of their time quarrelling, intriguing and splitting up their forces into ill-coordinated parts

To his disappointment, he failed to persuade the government to prosecute a major descent from the sea against Cadiz, but instead found himself landing at Alicante with his brother Michael as part of reinforcements for the Huguenot commander-in-chief, Lord Galway. Subsequently, as has already been related, he was appointed Governor of Alicante in succession to General Gorges, who returned to England

Recognizing that its defence could have only one outcome, and anxious to spare the townsfolk the fate of their compatriots at Denia where all had been put to the sword after the storm of the lower town, Richards beat a parley, and agreed to evacuate the town in return for an undertaking that the citizens should be treated by the Bourbon forces as if they had never rebelled against Philip V

On Monday, March 3rd, sentries reported at six in the morning that the townsfolk were hastily evacuating the quarter of Alicante nearest the castle

Again, the fact that he was not technically an English officer but a general in the service of Charles III, the Spanish Pretender, may have caused his act of courage to go relatively unrecognised in the history of a long war that saw many greater events than the siege of Alicante

On April 6, 1814, Napoleon, then in his mid-40s, was forced to abdicate the throne. With the Treaty of Fontainebleau, he was exiled to Elba, a Mediterranean island off the coast of Italy.

The new king, Louis XVIII, fled, and Napoleon embarked on what came to be known as his Hundred Days campaign.

Upon Napoleon’s return to France, a coalition of allies—the Austrians, British, Prussians and Russians—who considered the French emperor an enemy began to prepare for war. Napoleon raised a new army and planned to strike preemptively, defeating the allied forces one by one before they could launch a united attack against him.

Two days later, on June 18, Napoleon led his army of some 72,000 troops against the 68,000-man British army, which had taken up a position south of Brussels near the village of Waterloo.

Although Napoleon’s troops mounted a strong attack against the British, the arrival of the Prussians turned the tide against the French. The French emperor’s outnumbered army retreated in chaos.

Ultimately, the Battle of Waterloo marked the end of Napoleon’s storied military career. He reportedly rode away from the battle in tears.

Did you know? Today, the expression that someone has “met his Waterloo” means the person has suffered a decisive or final defeat or setback.

On June 22, 1815, Napoleon once again abdicated. That October, he was exiled to the remote, British-held island of Saint Helena, in the South Atlantic Ocean. He died there on May 5, 1821, at age 51, most likely from stomach cancer.

From the start, the British Navy far outmatched that of France. An oceanic superpower whose fleet dominated the seas, Britain repeatedly beat France. The most famous defeats were both delivered by Admiral Nelson, who destroyed Napoleon’s transport fleet at the Battle of the Nile and then smashed a Franco-Spanish force at Trafalgar, where the British Admiral lost his life.

As a mercantile nation, the British were reliant upon international trade. Napoleon, therefore, tried to win the economic war through the Continental System, a blockade of British trade at ports across Europe. It was easier to enforce in some areas than others. Anywhere directly controlled by France, Napoleon could order the system into place. Elsewhere, he had to win cooperation through diplomacy.

The system was full of holes. Occupied territories became resentful of the imposition, stirring opposition to the French. France’s Atlantic ports were hit hard by the British counter-blockade, their trade and supporting industries badly savaged. In the Netherlands, Napoleon’s brother failed to crack down on smuggling, while in Russia the Tsar gave up on the blockade as unhelpful for his nation.Not only did the Continental System fail to cripple Britain, it also damaged France’s own economy.

Between the driving determination of Tsar Alexander of Russia; the international influence of Austria; the freshly mobilized and experienced armies of Prussia; and the financial and military power of Great Britain, this alliance was finally able to push Napoleon’s troops back into France and defeat them there. He had offended too many people, leaving his nation without potential allies. The Coalition finally brought together enough strength to bring him down.

Napoleon always had to win on his terms. Ultimately, that led him to lose.