he first participants in the trade were elephants, the only group for whom ivory was truly essential. Tusks had and have important functions for elephants. They are used in feeding, in marking territory, as both offensive and defensive weapons, and as markers of status (Shoshani, 1992, p. 48). The questions for further study arising here relate to the ways in which hunting by humans affected elephant populations. To what extent were their physical reproduction and collective behaviour affected as they were reduced in numbers left: 263.997px; top: 561.245px; fon