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According to a paper in PLoS Computational Biology, "PageRank" can be applied to the study of food webs.

"In particular, rapid swings in the density of plants and fleas indicated the food web was unstable and about to change. The idea that such early warning signals ought to exist is not new - but the researchers say this is the first time it has been demonstrated experimentally. "For a long time, ecologists thought these changes couldn't be predicted," said research leader Stephen Carpenter from the University of Wisconsin in Madison, US. "But we've now shown that they can be foreseen. The early warning is clear; it is a strong signal.""