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Until now, research into regime shifts has focused on critical environmental thresholds, or "tipping points," in external conditions-eg when crossing a certain temperature threshold triggers a sudden shift to desertification. But the new model by Catalina Chaparro-Pedraza and SFI External Professor André de Roos, both at the University of Amsterdam, reveals how a small change in the external environment, with little immediate impact, can induce slow evolutionary changes in the species that inhabit the system. After what the researchers call a "considerable delay," wherein species slowly evolve a new trait or behavior over generations, the regime shift manifests as a delayed reaction.